The Evolution of Man, V.1., Ernst Haeckel [ebook pdf reader for pc .TXT] 📗

interesting lectures which made up this work he proved clearly that the descent of man from the ape followed necessarily from the theory of descent. If that theory is true, we are bound to conceive the animals which most closely resemble man as those from which humanity has been gradually evolved. About the same time Carl Vogt published a larger work on the same subject. We must also mention Gustav Jaeger and Friedrich Rolle among the zoologists who accepted and taught the theory of evolution immediately after the publication of Darwin's book, and maintained that the descent of man from the lower animals logically followed from it. The latter published, in 1866, a work on the origin and position of man.

About the same time I attempted, in the second volume of my General Morphology (1866), to apply the theory of evolution to the whole organic kingdom, including man.* (* Huxley spoke of this "as one of the greatest scientific works ever published."--Translator.) I endeavoured to sketch the probable ancestral trees of the various classes of the animal world, the protists, and the plants, as it seemed necessary to do on Darwinian principles, and as we can actually do now with a high degree of confidence. If the theory of descent, which Lamarck first clearly formulated and Darwin thoroughly established, is true, we should be able to draw up a natural classification of plants and animals in the light of their genealogy, and to conceive the large and small divisions of the system as the branches and twigs of an ancestral tree. The eight genealogical tables which I inserted in the second volume of the General Morphology are the first sketches of their kind. In

Chapter 2.

27, particularly, I trace the chief stages in man's ancestry, as far as it is possible to follow it through the vertebrate stem. I tried especially to determine, as well as one could at that time, the position of man in the classification of the mammals and its genealogical significance. I have greatly improved this attempt, and treated it in a more popular form, in chapters 26 to 28 of my History of Creation (1868).* (* Of which Darwin said that the Descent of Man would probably never have been written if he had seen it earlier.--Translator.)

It was not until 1871, twelve years after the appearance of The Origin of Species, that Darwin published the famous work which made the much-contested application of his theory to man, and crowned the splendid structure of his system. This important work was The Descent of Man, and Selection in Relation to Sex. In this Darwin expressly drew the conclusion, with rigorous logic, that man also must have been developed out of lower species, and described the important part played by sexual selection in the elevation of man and the other higher animals. He showed that the careful selection which the sexes exercise on each other in regard to sexual relations and procreation, and the aesthetic feeling which the higher animals develop through this, are of the utmost importance in the progressive development of forms and the differentiation of the sexes. The males choosing the handsomest females in one class of animals, and the females choosing only the finest-looking males in another, the special features and the sexual characteristics are increasingly accentuated. In fact, some of the higher animals develop in this connection a finer taste and judgment than man himself. But, even as regards man, it is to this sexual selection that we owe the family-life, which is the chief foundation of civilisation. The rise of the human race is due for the most part to the advanced sexual selection which our ancestors exercised in choosing their mates.

Darwin accepted in the main the general outlines of man's ancestral tree, as I gave it in the General Morphology and the History of Creation, and admitted that his studies led him to the same conclusion. That he did not at once apply the theory to man in his first work was a commendable piece of discretion; such a sequel was bound to excite the strongest opposition to the whole theory. The first thing to do was to establish it as regards the animal and plant worlds. The subsequent extension to man was bound to be made sooner or later.

It is important to understand this very clearly. If all living things come from a common root, man must be included in the general scheme of evolution. On the other hand, if the various species were separately created, man, too, must have been created, and not evolved. We have to choose between these two alternatives. This cannot be too frequently or too strongly emphasised. EITHER all the species of animals and plants are of supernatural origin--created, not evolved--and in that case man also is the outcome of a creative act, as religion teaches, OR the different species have been evolved from a few common, simple ancestral forms, and in that case man is the highest fruit of the tree of evolution.

We may state this briefly in the following principle--The descent of man from the lower animals is a special deduction which inevitably follows from the general inductive law of the whole theory of evolution. In this principle we have a clear and plain statement of the matter. Evolution is in reality nothing but a great induction, which we are compelled to make by the comparative study of the most important facts of morphology and physiology. But we must draw our conclusion according to the laws of induction, and not attempt to determine scientific truths by direct measurement and mathematical calculation. In the study of living things we can scarcely ever directly and fully, and with mathematical accuracy, determine the nature of phenomena, as is done in the simpler study of the inorganic world--in chemistry, physics, mineralogy, and astronomy. In the latter, especially, we can always use the simplest and absolutely safest method--that of mathematical determination. But in biology this is quite impossible for various reasons; one very obvious reason being that most of the facts of the science are very complicated and much too intricate to allow a direct mathematical analysis. The greater part of the phenomena that biology deals with are complicated HISTORICAL PROCESSES, which are related to a far-reaching past, and as a rule can only be approximately estimated. Hence we have to proceed by INDUCTION--that is to say, to draw general conclusions, stage by stage, and with proportionate confidence, from the accumulation of detailed observations. These inductive conclusions cannot command absolute confidence, like mathematical axioms; but they approach the truth, and gain increasing probability, in proportion as we extend the basis of observed facts on which we build. The importance of these inductive laws is not diminished from the circumstance that they are looked upon merely as temporary acquisitions of science, and may be improved to any extent in the progress of scientific knowledge. The same may be said of the attainments of many other sciences, such as geology or archeology. However much they may be altered and improved in detail in the course of time, these inductive truths may retain their substance unchanged.

Now, when we say that the theory of evolution in the sense of Lamarck and Darwin is an inductive law--in fact, the greatest of all biological inductions--we rely, in the first place, on the facts of paleontology. This science gives us some direct acquaintance with the historical phenomena of the changes of species. From the situations in which we find the fossils in the various strata of the earth we gather confidently, in the first place, that the living population of the earth has been gradually developed, as clearly as the earth's crust itself; and that, in the second place, several different populations have succeeded each other in the various geological periods. Modern geology teaches that the formation of the earth has been gradual, and unbroken by any violent revolutions. And when we compare together the various kinds of animals and plants which succeed each other in the history of our planet, we find, in the first place, a constant and gradual increase in the number of species from the earliest times until the present day; and, in the second place, we notice that the forms in each great group of animals and plants also constantly improve as the ages advance. Thus, of the vertebrates there are at first only the lower fishes; then come the higher fishes, and later the amphibia. Still later appear the three higher classes of vertebrates--the reptiles, birds, and mammals, for the first time; only the lowest and least perfect forms of the mammals are found at first; and it is only at a very late period that placental mammals appear, and man belongs to the latest and youngest branch of these. Thus perfection of form increases as well as variety from the earliest to the latest stage. That is a fact of the greatest importance. It can only be explained by the theory of evolution, with which it is in perfect harmony. If the different groups of plants and animals do really descend from each other, we must expect to find this increase in their number and perfection under the influence of natural selection, just as the succession of fossils actually discloses it to us.

Comparative anatomy furnishes a second series of facts which are of great importance for the forming of our inductive law. This branch of morphology compares the adult structures of living things, and seeks in the great variety of organic forms the stable and simple law of organisation, or the common type or structure. Since Cuvier founded this science at the beginning of the nineteenth century it has been a favourite study of the most distinguished scientists. Even before Cuvier's time Goethe had been greatly stimulated by it, and induced to take up the study of morphology. Comparative osteology, or the philosophic study and comparison of the bony skeleton of the vertebrates--one of its most interesting sections--especially fascinated him, and led him to form the theory of the skull which I mentioned before. Comparative anatomy shows that the internal structure of the animals of each stem and the plants of each class is the same in its essential features, however much they differ in external appearance. Thus man has so great a resemblance in the chief features of his internal organisation to the other mammals that no comparative anatomist has ever doubted that he belongs to this class. The whole internal structure of the human body, the arrangement of its various systems of organs, the distribution of the bones, muscles, blood-vessels, etc., and the whole structure of these organs in the larger and the finer scale, agree so closely with those of the other mammals (such as the apes, rodents, ungulates, cetacea, marsupials, etc.) that their external differences are of no account whatever. We learn further from comparative anatomy that the chief features of animal structure are so similar in the various classes (fifty to sixty in number altogether) that they may all be comprised in from eight to twelve great groups. But even in these groups, the stem-forms or animal types, certain organs (especially the alimentary canal) can be proved to have been originally the same for all. We can only explain by the theory of evolution this essential unity in internal structure of all these animal forms that differ so much in outward appearance. This wonderful fact can only be really understood and explained when we regard the internal resemblance as an inheritance from common-stem forms, and the external differences as the effect of adaptation to different environments.

In recognising this, comparative anatomy has itself advanced to a higher stage. Gegenbaur, the most distinguished of recent students of this science, says that with the theory of evolution a new period began in comparative anatomy, and that the theory in turn found a touch stone in the science. "Up to now there is no fact in comparative anatomy