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perfect gentleman, but so very dissatisfied with that vapid consolation that for hours he continues his imaginary rehearsals of what he would have liked to have done to that Custodian (mostly with his boots) by way of teaching him manners.

Some time after this I visited the Natural History Museum with Captain Scott’s sister. After a slight preliminary skirmish in which we convinced a minor custodian that the specimens brought by the expedition from the Antarctic did not include the moths we found preying on some of them, Miss Scott expressed a wish to see the penguins’ eggs. Thereupon the minor custodians flatly denied that any such eggs were in existence or in their possession. Now Miss Scott was her brother’s sister; and she showed so little disposition to take this lying down that I was glad to get her away with no worse consequences than a profanely emphasized threat on my part that if we did not receive ample satisfaction in writing within twenty-four hours as to the safety of the eggs England would reverberate with the tale.

The ultimatum was effectual; and due satisfaction was forthcoming in time; but I was relieved when I learnt later on that they had been entrusted to Professor Assheton for the necessary microscopic examination. But he died before he could approach the task; and the eggs passed into the hands of Professor Cossar Ewart of Edinburgh University.

His report is as follows:

Appendix Professor Cossar Ewart’s Report

“It was a great disappointment to Dr. Wilson that no Emperor Penguin embryos were obtained during the cruise of the Discovery. But though embryos were conspicuous by their absence in the Emperor eggs brought home by the National Antarctic Expedition, it is well to bear in mind that the naturalists on board the Discovery learned much about the breeding habits of the largest living member of the ancient penguin family. Amongst other things it was ascertained (1) that in the case of the Emperor, as in the King Penguin, the egg during the period of incubation rests on the upper surface of the feet protected and kept in position by a fold of skin from the lower breast; and (2) that in the case of the Emperor the whole process of incubation is carried out on sea ice during the coldest and darkest months of the antarctic winter.

“After devoting much time to the study of penguins Dr. Wilson came to the conclusion that Emperor embryos would throw new light on the origin and history of birds, and decided that if he again found his way to the Antarctic he would make a supreme effort to visit an Emperor rookery during the breeding season. When, and under what conditions, the Cape Crozier rookery was eventually visited and Emperor eggs secured is graphically told in The Winter Journey. The question now arises, Has ‘the weirdest bird’s-nesting expedition that has ever been made’ added appreciably to our knowledge of birds?

“It is admitted that birds are descended from bipedal reptiles which flourished some millions of years ago⁠—reptiles in build not unlike the kangaroo. From Archaeopteryx of Jurassic times we know primeval birds had teeth, three fingers with claws on each hand, and a long lizard-like tail provided with nearly twenty pairs of well-formed true feathers. But unfortunately neither this lizard-tailed bird, nor yet the fossil birds found in America, throw any light on the origin of feathers. Ornithologists and others who have devoted much time to the study of birds have as a rule assumed that feathers were made out of scales, that the scales along the margin of the hand and forearm and along each side of the tail were elongated, frayed and otherwise modified to form the wing and tail quills, and that later other scales were altered to provide a coat capable of preventing loss of heat. But as it happens, a study of the development of feathers affords no evidence that they were made out of scales. There are neither rudiments of scales nor feathers in very young bird embryos. In the youngest of the three Emperor embryos there are, however, feather rudiments in the tail region⁠—the embryo was probably seven or eight days old⁠—but in the two older embryos there are a countless number of feather rudiments, i.e. of minute pimples known as papillae.

“In penguins as in many other birds there are two distinct crops of feather papillae, viz.: a crop of relatively large papillae which develop into prepennae, the forerunners of true feathers (pennae), and a crop of small papillae which develop into preplumulae, the forerunners of true down feathers (plumulae).

“In considering the origin of feathers we are not concerned with the true feathers (pennae), but with the nestling feathers (prepennae), and more especially with the papillae from which the prepennae are developed. What we want to know is, Do the papillae which in birds develop into the first generation of feathers correspond to the papillae which in lizards develop into scales?

“The late Professor Assheton, who undertook the examination of some of the material brought home by the Terra Nova, made a special study of the feather papillae of the Emperor Penguin embryos from Cape Crozier. Drawings were made to indicate the number, size and time of appearance of the feather papillae, but unfortunately in the notes left by the distinguished embryologist there is no indication whether the feather papillae were regarded as modified scale papillae or new creations resulting from the appearance of special feather-forming factors in the germ-plasm.

“When eventually the three Emperor Penguin embryos reached me that their feather rudiments might be compared with the feather rudiments of other birds, I noticed that in Emperor embryos the feather papillae appeared before the scale papillae. Evidence of this was especially afforded by the largest embryo, which had reached about the same stage in its development as a 16-days goose embryo.

“In the largest Emperor embryo feather papillae occur all over the hindquarters and on the legs to within a short distance of the

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