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Mr. A. Murray has carefully examined the Pediculi collected in different countries from the different races of man (8. ‘Transactions of the Royal Society of Edinburgh,’ vol. xxii, 1861, p. 567.); and he finds that they differ, not only in colour, but in the structure of their claws and limbs. In every case in which many specimens were obtained the differences were constant. The surgeon of a whaling ship in the Pacific assured me that when the Pediculi, with which some Sandwich Islanders on board swarmed, strayed on to the bodies of the English sailors, they died in the course of three or four days. These Pediculi were darker coloured, and appeared different from those proper to the natives of Chiloe in South America, of which he gave me specimens. These, again, appeared larger and much softer than European lice. Mr. Murray procured four kinds from Africa, namely, from the Negroes of the Eastern and Western coasts, from the Hottentots and Kaffirs; two kinds from the natives of Australia; two from North and two from South America. In these latter cases it may be presumed that the Pediculi came from natives inhabiting different districts. With insects slight structural differences, if constant, are generally esteemed of specific value: and the fact of the races of man being infested by parasites, which appear to be specifically distinct, might fairly be urged as an argument that the races themselves ought to be classed as distinct species.

Our supposed naturalist having proceeded thus far in his investigation, would next enquire whether the races of men, when crossed, were in any degree sterile. He might consult the work (9. ‘On the Phenomena of Hybridity in the Genus Homo,’ Eng. translat., 1864.) of Professor Broca, a cautious and philosophical observer, and in this he would find good evidence that some races were quite fertile together, but evidence of an opposite nature in regard to other races. Thus it has been asserted that the native women of Australia and Tasmania rarely produce children to European men; the evidence, however, on this head has now been shewn to be almost valueless. The half-castes are killed by the pure blacks: and an account has lately been published of eleven half-caste youths murdered and burnt at the same time, whose remains were found by the police. (10. See the interesting letter by Mr. T.A. Murray, in the ‘Anthropological Review,’ April 1868, p. liii. In this letter Count Strzelecki’s statement that Australian women who have borne children to a white man, are afterwards sterile with their own race, is disproved. M. A. de Quatrefages has also collected (Revue des Cours Scientifiques, March, 1869, p. 239), much evidence that Australians and Europeans are not sterile when crossed.) Again, it has often been said that when mulattoes intermarry, they produce few children; on the other hand, Dr. Bachman, of Charleston (11. ‘An Examination of Prof. Agassiz’s Sketch of the Nat. Provinces of the Animal World,’ Charleston, 1855, p. 44.), positively asserts that he has known mulatto families which have intermarried for several generations, and have continued on an average as fertile as either pure whites or pure blacks. Enquiries formerly made by Sir C. Lyell on this subject led him, as he informs me, to the same conclusion. (12. Dr. Rohlfs writes to me that he found the mixed races in the Great Sahara, derived from Arabs, Berbers, and Negroes of three tribes, extraordinarily fertile. On the other hand, Mr. Winwood Reade informs me that the Negroes on the Gold Coast, though admiring white men and mulattoes, have a maxim that mulattoes should not intermarry, as the children are few and sickly. This belief, as Mr. Reade remarks, deserves attention, as white men have visited and resided on the Gold Coast for four hundred years, so that the natives have had ample time to gain knowledge through experience.) In the United States the census for the year 1854 included, according to Dr. Bachman, 405,751 mulattoes; and this number, considering all the circumstances of the case, seems small; but it may partly be accounted for by the degraded and anomalous position of the class, and by the profligacy of the women. A certain amount of absorption of mulattoes into negroes must always be in progress; and this would lead to an apparent diminution of the former. The inferior vitality of mulattoes is spoken of in a trustworthy work (13. ‘Military and Anthropological Statistics of American Soldiers,’ by B.A. Gould, 1869, p. 319.) as a well-known phenomenon; and this, although a different consideration from their lessened fertility, may perhaps be advanced as a proof of the specific distinctness of the parent races. No doubt both animal and vegetable hybrids, when produced from extremely distinct species, are liable to premature death; but the parents of mulattoes cannot be put under the category of extremely distinct species. The common Mule, so notorious for long life and vigour, and yet so sterile, shews how little necessary connection there is in hybrids between lessened fertility and vitality; other analogous cases could be cited.

Even if it should hereafter be proved that all the races of men were perfectly fertile together, he who was inclined from other reasons to rank them as distinct species, might with justice argue that fertility and sterility are not safe criterions of specific distinctness. We know that these qualities are easily affected by changed conditions of life, or by close inter-breeding, and that they are governed by highly complex laws, for instance, that of the unequal fertility of converse crosses between the same two species. With forms which must be ranked as undoubted species, a perfect series exists from those which are absolutely sterile when crossed, to those which are almost or completely fertile. The degrees of sterility do not coincide strictly with the degrees of difference between the parents in external structure or habits of life. Man in many respects may be compared with those animals which have long been domesticated, and a large body of evidence can be advanced in favour of the Pallasian doctrine (14. The ‘Variation of Animals and Plants under Domestication,’ vol. ii. p. 109. I may here remind the reader that the sterility of species when crossed is not a specially-acquired quality, but, like the incapacity of certain trees to be grafted together, is incidental on other acquired differences. The nature of these differences is unknown, but they relate more especially to the reproductive system, and much less so to external structure or to ordinary differences in constitution. One important element in the sterility of crossed species apparently lies in one or both having been long habituated to fixed conditions; for we know that changed conditions have a special influence on the reproductive system, and we have good reason to believe (as before remarked) that the fluctuating conditions of domestication tend to eliminate that sterility which is so general with species, in a natural state, when crossed. It has elsewhere been shewn by me (ibid. vol. ii. p. 185, and ‘Origin of Species,’ 5th edit. p. 317), that the sterility of crossed species has not been acquired through natural selection: we can see that when two forms have already been rendered very sterile, it is scarcely possible that their sterility should be augmented by the preservation or survival of the more and more sterile individuals; for, as the sterility increases, fewer and fewer offspring will be produced from which to breed, and at last only single individuals will be produced at the rarest intervals. But there is even a higher grade of sterility than this. Both Gartner and Kolreuter have proved that in genera of plants, including many species, a series can be formed from species which, when crossed, yield fewer and fewer seeds, to species which never produce a single seed, but yet are affected by the pollen of the other species, as shewn by the swelling of the germen. It is here manifestly impossible to select the more sterile individuals, which have already ceased to yield seeds; so that the acme of sterility, when the germen alone is affected, cannot have been gained through selection. This acme, and no doubt the other grades of sterility, are the incidental results of certain unknown differences in the constitution of the reproductive system of the species which are crossed.), that domestication tends to eliminate the sterility which is so general a result of the crossing of species in a state of nature. From these several considerations, it may be justly urged that the perfect fertility of the intercrossed races of man, if established, would not absolutely preclude us from ranking them as distinct species.

Independently of fertility, the characters presented by the offspring from a cross have been thought to indicate whether or not the parent-forms ought to be ranked as species or varieties; but after carefully studying the evidence, I have come to the conclusion that no general rules of this kind can be trusted. The ordinary result of a cross is the production of a blended or intermediate form; but in certain cases some of the offspring take closely after one parent-form, and some after the other. This is especially apt to occur when the parents differ in characters which first appeared as sudden variations or monstrosities. (15. ‘The Variation of Animals,’ etc., vol. ii. p. 92.) I refer to this point, because Dr. Rohlfs informs me that he has frequently seen in Africa the offspring of negroes crossed with members of other races, either completely black or completely white, or rarely piebald. On the other hand, it is notorious that in America mulattoes commonly present an intermediate appearance.

We have now seen that a naturalist might feel himself fully justified in ranking the races of man as distinct species; for he has found that they are distinguished by many differences in structure and constitution, some being of importance. These differences have, also, remained nearly constant for very long periods of time. Our naturalist will have been in some degree influenced by the enormous range of man, which is a great anomaly in the class of mammals, if mankind be viewed as a single species. He will have been struck with the distribution of the several so-called races, which accords with that of other undoubtedly distinct species of mammals. Finally, he might urge that the mutual fertility of all the races has not as yet been fully proved, and even if proved would not be an absolute proof of their specific identity.

On the other side of the question, if our supposed naturalist were to enquire whether the forms of man keep distinct like ordinary species, when mingled together in large numbers in the same country, he would immediately discover that this was by no means the case. In Brazil he would behold an immense mongrel population of Negroes and Portuguese; in Chiloe, and other parts of South America, he would behold the whole population consisting of Indians and Spaniards blended in various degrees. (16. M. de Quatrefages has given (‘Anthropological Review,’ Jan. 1869, p. 22), an interesting account of the success and energy of the Paulistas in Brazil, who are a much crossed race of Portuguese and Indians, with a mixture of the blood of other races.) In many parts of the same continent he would meet with the most complex crosses between Negroes, Indians, and Europeans; and judging from the vegetable kingdom, such triple crosses afford the severest test of the mutual fertility of the parent forms. In one island of the Pacific he would find a small population of mingled Polynesian and English blood; and in the Fiji Archipelago a

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