The Voyage of the Beagle, Charles Robert Darwin [book recommendations .txt] 📗

in which these remains were embedded, contains, according to Professor Ehrenberg, eight fresh-water and one salt-water infusorial animalcule; therefore, probably, it was an estuary deposit.
The remains at Punta Alta were embedded in stratified gravel and reddish mud, just such as the sea might now wash up on a shallow bank. They were associated with twenty- three species of shells, of which thirteen are recent and four others very closely related to recent forms. [1] From the bones of the Scelidotherium, including even the knee-cap, being intombed in their proper relative positions, and from the osseous armour of the great armadillo-like animal being so well preserved, together with the bones of one of its legs, we may feel assured that these remains were fresh and united by their ligaments, when deposited in the gravel together with the shells. [2] Hence we have good evidence that the above enumerated gigantic quadrupeds, more different from those of the present day than the oldest of the tertiary quadrupeds of Europe, lived whilst the sea was peopled with most of its present inhabitants; and we have confirmed that remarkable law so often insisted on by Mr. Lyell, namely, that the "longevity of the species in the mammalia is upon the whole inferior to that of the testacea." [3]
The great size of the bones of the Megatheroid animals, including the Megatherium, Megalonyx, Scelidotherium, and Mylodon, is truly wonderful. The habits of life of these animals were a complete puzzle to naturalists, until Professor Owen [4] solved the problem with remarkable ingenuity. The teeth indicate, by their simple structure, that these Megatheroid animals lived on vegetable food, and probably on the leaves and small twigs of trees; their ponderous forms and great strong curved claws seem so little adapted for locomotion, that some eminent naturalists have actually believed, that, like the sloths, to which they are intimately related, they subsisted by climbing back downwards on trees, and feeding on the leaves. It was a bold, not to say preposterous, idea to conceive even antediluvian trees, with branches strong enough to bear animals as large as elephants. Professor Owen, with far more probability, believes that, instead of climbing on the trees, they pulled the branches down to them, and tore up the smaller ones by the roots, and so fed on the leaves. The colossal breadth and weight of their hinder quarters, which can hardly be imagined without having been seen, become on this view, of obvious service, instead of being an incumbrance: their apparent clumsiness disappears. With their great tails and their huge heels firmly fixed like a tripod on the ground, they could freely exert the full force of their most powerful arms and great claws. Strongly rooted, indeed, must that tree have been, which could have resisted such force! The Mylodon, moreover, was furnished with a long extensile tongue like that of the giraffe, which, by one of those beautiful provisions of nature, thus reaches with the aid of its long neck its leafy food. I may remark, that in Abyssinia the elephant, according to Bruce, when it cannot reach with its proboscis the branches, deeply scores with its tusks the trunk of the tree, up and down and all round, till it is sufficiently weakened to be broken down.
The beds including the above fossil remains, stand only from fifteen to twenty feet above the level of high-water; and hence the elevation of the land has been small (without there has been an intercalated period of subsidence, of which we have no evidence) since the great quadrupeds wandered over the surrounding plains; and the external features of the country must then have been very nearly the same as now. What, it may naturally be asked, was the character of the vegetation at that period; was the country as wretchedly sterile as it now is? As so many of the co-embedded shells are the same with those now living in the bay, I was at first inclined to think that the former vegetation was probably similar to the existing one; but this would have been an erroneous inference for some of these same shells live on the luxuriant coast of Brazil; and generally, the character of the inhabitants of the sea are useless as guides to judge of those on the land. Nevertheless, from the following considerations, I do not believe that the simple fact of many gigantic quadrupeds having lived on the plains round Bahia Blanca, is any sure guide that they formerly were clothed with a luxuriant vegetation: I have no doubt that the sterile country a little southward, near the Rio Negro, with its scattered thorny trees, would support many and large quadrupeds.
That large animals require a luxuriant vegetation, has been a general assumption which has passed from one work to another; but I do not hesitate to say that it is completely false, and that it has vitiated the reasoning of geologists on some points of great interest in the ancient history of the world. The prejudice has probably been derived from India, and the Indian islands, where troops of elephants, noble forests, and impenetrable jungles, are associated together in every one's mind. If, however, we refer to any work of travels through the southern parts of Africa, we shall find allusions in almost every page either to the desert character of the country, or to the numbers of large animals inhabiting it. The same thing is rendered evident by the many engravings which have been published of various parts of the interior. When the Beagle was at Cape Town, I made an excursion of some days' length into the country, which at least was sufficient to render that which I had read more fully intelligible.
Dr. Andrew Smith, who, at the head of his adventurous party, has lately succeeded in passing the Tropic of Capricorn, informs me that, taking into consideration the whole of the southern part of Africa, there can be no doubt of its being a sterile country. On the southern and south-eastern coasts there are some fine forests, but with these exceptions, the traveller may pass for days together through open plains, covered by a poor and scanty vegetation. It is difficult to convey any accurate idea of degrees of comparative fertility; but it may be safely said that the amount of vegetation supported at any one time [5] by Great Britain, exceeds, perhaps even tenfold, the quantity on an equal area, in the interior parts of Southern Africa. The fact that bullock- waggons can travel in any direction, excepting near the coast, without more than occasionally half an hour's delay in cutting down bushes, gives, perhaps, a more definite notion of the scantiness of the vegetation. Now, if we look to the animals inhabiting these wide plains, we shall find their numbers extraordinarily great, and their bulk immense. We must enumerate the elephant, three species of rhinoceros, and probably, according to Dr. Smith, two others, the hippopotamus, the giraffe, the bos caffer -- as large as a full-grown bull, and the elan -- but little less, two zebras, and the quaccha, two gnus, and several antelopes even larger than these latter animals. It may be supposed that although the species are numerous, the individuals of each kind are few. By the kindness of Dr. Smith, I am enabled to show that the case is very different. He informs me, that in lat. 24 degs., in one day's march with the bullock-waggons, he saw, without wandering to any great distance on either side, between one hundred and one hundred and fifty rhinoceroses, which belonged to three species: the same day he saw several herds of giraffes, amounting together to nearly a hundred; and that although no elephant was observed, yet they are found in this district. At the distance of a little more than one hour's march from their place of encampment on the previous night, his party actually killed at one spot eight hippopotamuses, and saw many more. In this same river there were likewise crocodiles. Of course it was a case quite extraordinary, to see so many great animals crowded together, but it evidently proves that they must exist in great numbers. Dr. Smith describes the country passed through that day, as "being thinly covered with grass, and bushes about four feet high, and still more thinly with mimosa-trees." The waggons were not prevented travelling in a nearly straight line.
Besides these large animals, every one the least acquainted with the natural history of the Cape, has read of the herds of antelopes, which can be compared only with the flocks of migratory birds. The numbers indeed of the lion, panther, and hyaena, and the multitude of birds of prey, plainly speak of the abundance of the smaller quadrupeds: one evening seven lions were counted at the same time prowling round Dr. Smith's encampment. As this able naturalist remarked to me, the carnage each day in Southern Africa must indeed be terrific! I confess it is truly surprising how such a number of animals can find support in a country producing so little food. The larger quadrupeds no doubt roam over wide tracts in search of it; and their food chiefly consists of underwood, which probably contains much nutriment in a small bulk. Dr. Smith also informs me that the vegetation has a rapid growth; no sooner is a part consumed, than its place is supplied by a fresh stock. There can be no doubt, however, that our ideas respecting the apparent amount of food necessary for the support of large quadrupeds are much exaggerated: it should have been remembered that the camel, an animal of no mean bulk, has always been considered as the emblem of the desert.
The belief that where large quadrupeds exist, the vegetation must necessarily be luxuriant, is the more remarkable, because the converse is far from true. Mr. Burchell observed to me that when entering Brazil, nothing struck him more forcibly than the splendour of the South American vegetation contrasted with that of South Africa, together with the absence of all large quadrupeds. In his Travels, [6] he has suggested that the comparison of the respective weights (if there were sufficient data) of an equal number of the largest herbivorous quadrupeds of each country would be extremely curious. If we take on the one side, the elephant, [7] hippopotamus, giraffe, bos caffer, elan, certainly three, and probably five species of rhinoceros; and on the American side, two tapirs, the guanaco, three deer, the vicuna, peccari, capybara (after which we must choose from the monkeys to complete the number), and then place these two groups alongside each other, it is not easy to conceive ranks more disproportionate in size. After the above facts, we are compelled to conclude, against anterior probability, [8] that among the mammalia there exists no close relation between the bulk of the species, and the _quantity_ of the vegetation, in the countries which they inhabit.
With regard to the number of large quadrupeds, there certainly exists no quarter of the globe which will bear comparison with Southern Africa. After the different statements which have been given, the extremely desert character of that region will not be disputed. In the European division of the world, we must look back to the tertiary epochs, to find a condition of things among the mammalia, resembling that now existing at the Cape of Good Hope. Those tertiary epochs, which we are apt to consider as abounding to an astonishing degree with large animals, because we find the remains of many ages accumulated at certain spots, could hardly boast of more large quadrupeds than Southern Africa does at present. If we speculate on the condition of the vegetation